Actinopteri (ray-finned fishes) > Blenniiformes
(Blennies) > Tripterygiidae
(Triplefin blennies) > Tripterygiinae
Etymology: Helcogramma: Greek, helkos, -eos, -ous = ulcer, sore + Greek, gramma = letter, mark (Ref. 45335). More on authors: Fricke & Randall.
Environment: milieu / climate zone / depth range / distribution range
Marine; demersal; depth range - 10 m (Ref. 12235), usually - m (Ref. ). Tropical; 0°C -
Western Indian Ocean: presently endemic to the Maldives.
Size / Weight / Age
Maturity: Lm ?  range ? - ? cm
Max length : 2.8 cm SL male/unsexed; (Ref. 12235)
(total): 15 - 17;
soft rays: 17 - 21;
Vertebrae: 35 - 36. Diagnosis: Dorsal fin III + XIII+ 10, 2nd fin twice height of 1st; anal fin I + 16-18; pectoral rays 15 (3+6+6); pelvic fin I, 2, spine short and hidden, segmented rays simple, partially united by membrane. Vertebrae 10 + 25-26. LL total lateral scales 36-38; tubed 13-21 (16-17) scales, ends below 3rd dorsal fin; nape, belly, 1st dorsal fin and anterior anal-fin base scaleless, caudal-fin base with 2 rows of scales. Mandibular pores 3+2-1+3. Labial folds and orbital cirrus absent. Head length 3.4 in SL; eye 2.4, maxilla 2.3 in HL. Males overall reddish with bluish dots and streaks on head, forming stripes anteriorly on body, ventrally light grey, fins pigmented; females paler, without spots and stripes, ventrally white, fins transluscent (Ref. 75031).
Eggs are hemispherical and covered with numerous sticky threads that anchor them in the algae on the nesting sites (Ref. 240). Larvae are planktonic which occur primarily in shallow, nearshore waters (Ref. 94114).
Life cycle and mating behavior
Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae
Fricke, R. and J.E. Randall, 1992. Tripterygiid fishes of the Maldives Islands, with descriptions of two new species (Teleostei: Blennioidei). Stuttg. Beitr. Naturkd. A Biol. (484):1-13. (Ref. 12235)
IUCN Red List Status (Ref. 124695)
Threat to humans
Common namesSynonymsMetabolismPredatorsEcotoxicologyReproductionMaturitySpawningSpawning aggregationFecundityEggsEgg development
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingNutrientsMass conversion
Estimates based on models
Preferred temperature (Ref. 115969
): 28.4 - 29, mean 28.9 °C (based on 53 cells).
Phylogenetic diversity index (Ref. 82805
= 0.5000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00550 (0.00248 - 0.01216), b=3.08 (2.89 - 3.27), in cm total length, based on LWR estimates for this (Sub)family-body shape (Ref. 93245
Trophic level (Ref. 69278
): 3.0 ±0.3 se; based on size and trophs of closest relatives
Resilience (Ref. 120179
): High, minimum population doubling time less than 15 months (Preliminary K or Fecundity.).
Vulnerability (Ref. 59153
): Low vulnerability (10 of 100) .
Nutrients (Ref. 124155
): Calcium = 1140 [486, 2,589] mg/100g ; Iron = 2.36 [1.27, 4.84] mg/100g ; Protein = 17.8 [16.6, 19.0] % ; Omega3 = 0.278 [0.122, 0.635] g/100g ; Selenium = 43.4 [17.0, 108.8] μg/100g ; VitaminA = 41.4 [13.4, 138.3] μg/100g ; Zinc = 2.2 [1.3, 3.6] mg/100g (wet weight);