Environment: milieu / climate zone / depth range / distribution range
Marine; reef-associated; non-migratory; depth range 5 - 135 m (Ref. 90102), usually 5 - 20 m (Ref. 58302). Tropical; 30°N - 30°S
Indo-Pacific: East Africa to the Galapagos Islands, north to Ryukyu and Hawaiian islands, south to Australia and the Tuamoto Islands.
Size / Weight / Age
Maturity: Lm ?  range ? - ? cm
Max length : 10.0 cm TL male/unsexed; (Ref. 4313)
soft rays: 5 - 6. Tan to reddish or brown in color (Ref. 4313). Has prickly papillae instead of scales. Dorsal fin high, 3rd or 4th spine longest; suborbital ridge without spines or with lump at head of ridge; preopercle with 2 indistinct spines only; body extremely compressed; soft dorsal fin attached to the caudal fin; coloration is variable, from nearly all yellow to red, brown or nearly black and variously mottled with darker pigment (Ref. 10482).
Inhabits reef flats, outer reef slopes, current-swept channels, and rarely on lagoon reefs (Ref. 1602). Benthic (Ref. 58302). Solitary and usually immobile among algae or seagrass but effects hip movements resembling that of a leaf falling down from a tree. Molts twice a month with the skin breaking off first in the head region. Has the habit of mimicking a dead leaf by swaying from side to side (Ref. 37816). Feeds on small crustaceans and fishes (Ref. 9710); also feeds on larvae (Ref. 5503). Venomous spines.
Life cycle and mating behavior
Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae
Myers, R.F., 1991. Micronesian reef fishes. Second Ed. Coral Graphics, Barrigada, Guam. 298 p. (Ref. 1602)
IUCN Red List Status (Ref. 125652)
Common namesSynonymsMetabolismPredatorsEcotoxicologyReproductionMaturitySpawningSpawning aggregationFecundityEggsEgg development
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingNutrientsMass conversion
Estimates based on models
Preferred temperature (Ref. 123201
): 24.6 - 28.8, mean 27.5 °C (based on 566 cells).
Phylogenetic diversity index (Ref. 82804
= 1.0000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.01259 (0.00612 - 0.02592), b=3.03 (2.86 - 3.20), in cm total length, based on LWR estimates for this (Sub)family-body shape (Ref. 93245
Trophic level (Ref. 69278
): 4.1 ±0.68 se; based on food items.
Resilience (Ref. 120179
): High, minimum population doubling time less than 15 months (Preliminary K or Fecundity.).
Fishing Vulnerability (Ref. 59153
): Low vulnerability (10 of 100).
Nutrients (Ref. 124155
): Calcium = 116 [56, 321] mg/100g ; Iron = 0.881 [0.455, 1.653] mg/100g ; Protein = 18.2 [16.1, 20.1] % ; Omega3 = 0.274 [0.118, 0.750] g/100g ; Selenium = 27.2 [11.3, 64.3] μg/100g ; VitaminA = 328 [115, 899] μg/100g ; Zinc = 1.43 [0.91, 2.21] mg/100g (wet weight);