Snout rounded, but not swollen or projecting; upper jaw length 38-44 % of head length, lower jaw length 44-53 % of head length. First pair of mental pores close together, united by a crescent-shaped groove just behind symphysis. Teeth differentiated into large and small in both jaws. Second anal spine long, stiff, 39-59 % of head length. From 8 to 13 scale rows above lateral line to origin of dorsal fin, 12-17 scale rows below lateral line to origin of anal fin (total 21-29 scale rows). Swim bladder carrot-shaped, with 18-22 pairs of arborescent appendages along its sides, the first pair entering head beyond transverse septum. Sagitta (large earstone) with a tadpole-shaped impression, the tail of which is J-shaped.