This species is characterized by the following: usual number of vertebrae, precaudal 14 (rarely 13), caudal 53-54 (55), total 81-82 (80-84); number of fin rays, dorsal 123-131, anal 103-109, pectoral usually 20-21 (19-24); pyloric caeca usually 6 (5-7) in 2 whorls, usually 5-6+1; first gill arch with rudimentary rakers 4 (3-6) on the upper arm, 8 (7-10) developed rakers on the lower limb, 5-9 rudimentary rakers anteriorly on the first arch; total rakers 16-19; dorsal-fin origin often between neural spines 4-5 (11 specimens) and rarely between 2-3 or 3-4 (1 specimen) (Ref. 91765).