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|Sound produced||yes, active sound production|
|Type of sound produced||knocks (very weak)|
|Sound production organ||swim bladder (possibly opercula)|
|Sound mechanism||vibration by contraction of extrinsic muscles associated with swim bladder lobes, reinforced by general contraction of skeletal muscles or violent closing of opercula|
|Behavioural context||under duress (handling or moderate irritaion)|
|Reference||Fish, M.P. and W.H. Mowbray, 1970|
|Remark||recorded with highpass filter: 30 Hz|