| 主要参考文献 | Maitland, P.S. and R.N. Campbell, 1992 | ||
| 掠食阶段 | juv./adults | ||
| 食物一 | zoobenthos | ||
| 食物二 | worms | ||
| 食物三 | non-annelids | ||
| 食物群 | unidentified | 捕食阶段/部份 | juv./adults |
| 食物名称 | unidentified | ||
| 普通 | rare (1-5%) | ||
| 国家 | UK Scotland | ||
| 附注 | Locality: Altquhur Burn; no. of fish examined=255. Also Ref. 57821. | ||