| Main Ref. | Koli, L., 1990 |
| Mode | dioecism |
| Fertilization | external |
| Mating type | |
| Spawning frequency | one clear seasonal peak per year |
| Spawning aggregation | Ref. |
| Batch spawner | Yes. Ref. Kottelat, M. and J. Freyhof, 2007 |
| Reproductive guild |
nonguarders open water/substratum egg scatterers |
| Parental Care | none |
| Description of life cycle and mating behavior | Males and females form school separately and arrive on the spawning site at differen times. Spawn over clean gravel areas in flowing water or on wave-washed shores of lakes. Deposit eggs in shallow water on sand or gravel bottom which hatch in 4-5 days at 18°C. Females lay sticky eggs deep into clean gravel (Ref. 59043). |
| Search for more references on reproduction | Scirus |