|Coad, B.W., 1995
|once in a lifetime
|Description of life cycle and mating behavior
|Spawning occurs both daytime and nighttime. The female selects a site and builds a redd while the male courts her and drives away other males. Upon completion of the redd, the female settles into the pitt, immediately followed by the male. The pair releases eggs and sperm simultaneously and the eggs fall into the spaces between the gravel. The female then moves to the upstream edge of the redd and resumes digging, thereby covering the eggs. When completely covered, the female may use the new redd to spawn once more or she may look for another spot and repeat the entire process. Both males and females spawn with one or more members of the opposite sex (Ref. 12204, 27547). Reproductive strategy: synchronous ovarian organization, determinate fecundity (Ref. 51846).
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