| Citation | Niwa, T., 1987, Comparison of the gene frequency between sympatric populations of ninespine sticklebacks, genus Pungitius, in Hokkaido, Japan.. Jap. J. Ichthyol. 34(2):184-190. |
| Ref. | 12605 Niwa, T., 1987 |
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| Named Used as Valid | Pungitius pungitius |
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