Distinguishing characteristics: esca with 2 bulbous, narrowly based, distal appendages, anterior-most appendage approximately two-thirds to one-half length of posterior most, each covered with small digitiform papillae and clusters of tiny filaments around base; absence of bulbous distal prolongation; large, anteroposteriorly compressed, posterior escal appendage, divided along its distal margin into 3 or more lobes or several to many slender filaments; absence of posterolateral filaments; total number of teeth in upper jaw 21-42, lower jaw with 25-45 teeth; vomerine teeth 4-8; length of illicium 7.8-13.3% SL; head length 25.7-31.8; head width 14.5-19.7; head depth 26.2-32.7; length of premaxilla 17.6-21.6; length of lower jaw 21.8-31.0 (Ref. 86949).