Diagnosis: fronto-parietal fontanel absent in adults, sometimes pore-like in juveniles; snout short, more than 3x head lenght; dorsal fin origin at same level as, or just behind, pelvic fin insertion; sexual dimorphism affecting anal fin shape; head length/snout length 3.1-3.7; 24-30 lateral line scales; 4.5 scales between lateral line and dorsal fin; 9.5-11 predorsal scales; 11-13 anal fin branched rays; 16-21 gill rakers on lower limb of first gill arch; 8 teeth in outer premaxillary row; adults medium-sized; humeral spot always present; broad black band on flanks; most apparent part of this band not reaching or extending beyond vertical through posterior base of dorsal fin (Ref. 80290).

Description: fronto-parietal fontanel present in very small individuals, disappearing with growth; gill rakers rather thin and short; 8 outer premaxillary teeth aligned in a single row; anal fin with 3 branched and 11-13 unbranched rays (Ref. 80290). Adult males with marked sexual dimorphism of anal fin, more rounded than in females where fin is more concave distally (Ref. 28013).

Coloration: prominent humeral spot on third post-opercular scale; distinct longitudinal band from middle of rayed dorsal fin to adipose fin (situated dorsal to median rays of caudal fin); band is wider anteriorly than posteriorly, anteriorly it is about 1 scale wide; in preserved material fins generally greyish; in life Ref. 41597 writes: “fins sometimes yellow, sometimes orange; adipose fin sometimes bright red” (Ref. 80290).