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| Sound produced | yes, active sound production |
| Type of sound produced | knocks |
| Sound production organ | swim bladder |
| Sound mechanism | not specified |
| Behavioural context | when alarmed |
| Reference | Fish, M.P. and W.H. Mowbray, 1970 |
| Remark | recorded with bandpass filter: 30 - 1200 Hz, recording amplified by 6 dB, noise reduction filter applied |