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|Sound produced||yes, active sound production|
|Type of sound produced||toothy clicks & grunts|
|Sound production organ||pharyngeal teeth & swim bladder (similar to Haemulon album)|
|Sound mechanism||pharyngeal teeth stridulation amplified by adjacent swim bladder (similar to Haemulon album)|
|Behavioural context||under duress (manual stimulation)|
|Reference||Fish, M.P. and W.H. Mowbray, 1970|