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|Sound produced||yes, active sound production|
|Type of sound produced||scratchy grunts|
|Sound production organ||pharyngeal teeth & swim bladder|
|Sound mechanism||not specified, but probably pharyngeal teeth stridulation amplified by adjacent swim bladder|
|Behavioural context||under duress (manual stimulation) - easily induced sound production|
|Reference||Fish, M.P. and W.H. Mowbray, 1970|