| 主要參考文獻 | Oscoz, J., P.M. Leunda, R. Miranda and M.C. Escala, 2006 | ||
| 掠食階段 | adults | ||
| 食物一 | zoobenthos | ||
| 食物二 | insects | ||
| 食物三 | insects | ||
| 食物群 | Elmidae | 捕食階段/部份 | adults |
| 食物名稱 | water beetles | ||
| 普通 | 6.25 | common (6-20%) | |
| 國家 | Spain | ||
| 附註 | Locality: River Larraun, a tributary of Ebro River, Navarre, northern Spain. Fish length: >8 cm. | ||