Raja clavata, Thornback ray : fisheries, gamefish

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Raja clavata Linnaeus, 1758

Thornback ray
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Raja clavata   AquaMaps   Data sources: GBIF OBIS
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Image of Raja clavata (Thornback ray)
Raja clavata
Picture by Stergiou, K.I.

Classification / Names Noms communs | Synonymes | Catalog of Fishes (gen., sp.) | ITIS | CoL | WoRMS | Cloffa

Élasmobranches (requins et raies) (sharks and rays) > Rajiformes (Skates and rays) > Rajidae (Skates) > Rajinae
Etymology: Raja: Latin, raja, -ae = a sting ray (Raja sp.) (Ref. 45335).  More on author: Linnaeus.

Environment: milieu / climate zone / depth range / distribution range Écologie

; marin dmersal; profondeur 5 - 1020 m (Ref. 114953).   Subtropical; 70°N - 29°S, 25°W - 42°E

Distribution Pays | Zones FAO | Écosystèmes | Occurrences | Point map | Introductions | Faunafri

Eastern Atlantic and Southwest Indian Ocean: Iceland to Madagascar, including the Mediterranean Sea.

Length at first maturity / Taille / Poids / Âge

Maturity: Lm 76.6, range 47 - 87.5 cm
Max length : 105 cm TL mâle / non sexé; (Ref. 53748); 139.0 cm TL (female); common length : 85.0 cm TL mâle / non sexé; (Ref. 9987); poids max. publié: 18.0 kg (Ref. 35388); âge max. reporté: 15 années (Ref. 41305)

Description synthétique Morphologie | Morphométrie

Épines dorsales (Total): 0; Épines anales 0; Rayons mous anaux: 0. Sub-rhomboid in shape with pointed wing-tips; disc-width 1,25 to 1,36 times in its length, its length 1,70 to 1,83 times in total length. Short rostrum, rounded at his extremity; pectoral fins with clear angles on lateral side; triangular pelvic fins (Ref. 39246). Dorsally prickly; large females also prickly throughout their ventral surface; juveniles and large males prickly along the margins of their discs and the underside of their snout. 30-50 thorns form a median row from the nape to the first dorsal fin; additional large 'buckler' thorns with swollen bases scattered on upper surface of disc in adults (Ref. 3167). Coloration is highly variable, with the dorsal surface having all shades of brown with dark and light spots and blotches; ventral surface white (Ref. 3167, 58137).

Biologie     Glossaire (ex. epibenthic)

Inhabits shelf and upper slope waters. Reported depth ranges varies, from 10-300 m (Ref. 4426, 82319, 88187), and from 300-577 m in the eastern Ionian Sea (Ref. 56504); most common in coastal waters between 10-60 m depth (Ref. 58137). Tolerates low salinities (Ref. 88171). Found on mud, sand and gravel bottoms, rarely on rougher bottoms (Ref. 6808, 58137). Nocturnal species. Feeds on all kinds of bottom animals, preferably crustaceans (Ref. 3167) and fishes (Ref. 114953). Undertakes migrations with mean distances of 54-117 km per month; shows a clear annual migration cycle (Ref. 89017, 89018), moves from deeper offshore waters (10-30 m) in autumn and winter to shallower areas (<10 m) in spring (Ref. 82319). Young are non-migratory, inhabiting inshore nursery grounds (Ref. 89019, 89020, 89022); in the Bay of Douarnenez (France) they are found to remain in shallow waters for at least 2 years (Ref. 89023). Maturity age 7-8 years (Ref. 114953). Oviparous. Young may tend to follow large objects, such as their mother (Ref. 205). Detects weak electric fields generated by other organisms (prey detection and predator avoidance) but may also generate its own weak electric fields (Ref. 10311, 88171). Utilized fresh and frozen (Ref. 9987).

Life cycle and mating behavior Maturité | Reproduction | Frai | Œufs | Fécondité | Larves

Oviparous. Distinct pairing with embrace. Polyandrous species (Ref. 89024). Paired eggs are laid and deposited on shallow sand, mud, pebble or gravel bottoms (Ref. 205, 89025). Up to 170 egg cases can be laid by a single female in a year (Ref. 3167), but average fecundity is much lower (around 48-74 eggs) (Ref. 3603, 31302). In northwestern Europe, egg cases are laid during spring (Ref. 3167) and in the Mediterranean during winter and spring (Ref. 3167). Egg cases are oblong capsules with stiff pointed horns at the corners, each containing one embryo (Ref. 205). Capsules are 5.0-9.0 cm long without the horns (Ref. 41250, 88187) and 3.4-6.8 cm wide (Ref. 41250). Egg cases are anchored with an adhesive film (Ref. 82399). Embryos feed solely on yolk (Ref. 50449). Egg cases hatch after about 4-5 months and pups are about 11-13 cm TL (Ref. 88864). Mating season from February to September, peaking in June (Ref. 74501). Adults observed to form same-sex aggregations during the mating season with females moving to shallower inshore waters approximately a month before the males (Ref 3603, 58137, 74501). Mating does not occur in the Baltic Sea (Ref. 82311).

Référence principale Upload your references | Références | Coordinateur : McEachran, John | Collaborateurs

McEachran, J.D. and K.A. Dunn, 1998. Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia 1998(2):271-290. (Ref. 27314)

Statut dans la liste rouge de l'IUCN (Ref. 120744)

  Quasi-menacé (NT) ; Date assessed: 03 October 2005

CITES (Ref. 118484)

Not Evaluated

CMS (Ref. 116361)

Not Evaluated

Menace pour l'homme

  Harmless





Utilisations par l'homme

Pêcheries: commercial; pêche sportive: oui
FAO(pêcheries: production; publication : search) | FishSource | Sea Around Us

Plus d'informations

Noms communs
Synonymes
Métabolisme
Prédateurs
Écotoxicologie
Reproduction
Maturité
Frai
Rassemblement de ponte
Fécondité
Œufs
Développement de l'œuf
Références
Aquaculture
Profil d'aquaculture
Souches
Génétique
Fréquences alléliques
Héritabilité
Pathologies
Traitement
Mass conversion
Collaborateurs
Images
Stamps, Coins Misc.
Sons
Ciguatera
Vitesse
Type de nage
Surface branchiale
Otolithes
Cerveaux
Vision

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Estimates based on models

Preferred temperature (Ref. 115969): 7.1 - 15.8, mean 10.7 (based on 474 cells).
Phylogenetic diversity index (Ref. 82805):  PD50 = 0.5000   [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00240 (0.00202 - 0.00285), b=3.24 (3.20 - 3.28), in cm Total Length, based on LWR estimates for this species (Ref. 93245).
Niveau trophique (Ref. 69278):  3.8   ±0.2 se; Based on diet studies.
Generation time: 10.0 (6.1 - 11.8) years. Estimated as median LN(3)/K based on 18 growth studies.
Résilience (Ref. 120179):  Faible, temps minimum de doublement de population : 4,5 à 14 années (K=0.09-0.14; tm=10; tmax=23; Fec=150).
Prior r = 0.18, 95% CL = 0.07 - 0.45, Based on 4 stock assessments.
Vulnérabilité (Ref. 59153):  High to very high vulnerability (73 of 100) .
Catégorie de prix (Ref. 80766):   Medium.