Actinopterygii (ray-finned fishes) > Ophidiiformes
(Cusk eels) > Bythitidae
(Viviparous brotulas) > Bythitinae
Etymology: Grammonus: Greek, gramma = signal, mark + Greek, onos = hake. More on authors: Smith & Radcliffe.
Environment: milieu / climate zone / depth range / distribution range
Marine; bathydemersal; depth range 45 - 345 m (Ref. 34024). Deep-water
Indo-West Pacific: Red Sea, Japan, East China Sea, and the Philippines.
Size / Weight / Age
Maturity: Lm ?  range ? - ? cm
Max length : 21.0 cm SL male/unsexed; (Ref. 559)
Morphology | Morphometrics
Uncommon species found on soft bottom (Ref. 34024). Ref. 9942 gives maximum depth to be 1,260 m.
Life cycle and mating behavior
Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae
Nielsen, J.G., D.M. Cohen, D.F. Markle and C.R. Robins, 1999. Ophidiiform fishes of the world (Order Ophidiiformes). An annotated and illustrated catalogue of pearlfishes, cusk-eels, brotulas and other ophidiiform fishes known to date. FAO Fish. Synop. 125(18):178p. Rome: FAO. (Ref. 34024)
IUCN Red List Status (Ref. 120744)
CITES (Ref. 115941)
Threat to humans
Fisheries: of no interest
Common namesSynonymsMetabolismPredatorsEcotoxicologyReproductionMaturitySpawningSpawning aggregationFecundityEggsEgg development
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingMass conversion
Estimates of some properties based on models
Preferred temperature (Ref. 115969
): 12.9 - 26.5, mean 19.5 (based on 337 cells).
Phylogenetic diversity index (Ref. 82805
= 0.5005 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00389 (0.00180 - 0.00842), b=3.12 (2.94 - 3.30), in cm Total Length, based on all LWR estimates for this body shape (Ref. 93245
Trophic Level (Ref. 69278
): 3.6 ±0.5 se; Based on size and trophs of closest relatives
Resilience (Ref. 69278
): Medium, minimum population doubling time 1.4 - 4.4 years (Preliminary K or Fecundity.).
Vulnerability (Ref. 59153
): Low to moderate vulnerability (34 of 100) .