Africa: Nilo-Sudanese species, known from the Nile basin and in West Africa from the Senegal River eastwards to the Cross, Wouri and Sanaga River, including the Chad basin (Ref. 43912). Various reports, including from the Ogowe and Congo River basin or Zambia and Zimbabwe, are based on misidentifications (Ref. 43912). Reports from Angola and Tanzania are questionable and need confirmation.