| Citation | Bussing, W.A., 1998, Peces de las aguas continentales de Costa Rica [Freshwater fishes of Costa Rica]. 2nd Ed.. San José Costa Rica: Editorial de la Universidad de Costa Rica. 468 p. |
| Ref. | 36880 Bussing, W.A., 1998 |
| Page | 332-334 |
| Named Used as Valid | Astatheros rhytisma |
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